This exhibition invites visitors to discover an example of connection between nature and culture. Through a richly evocative journey, it will illustrate how cooperation among people is the driving force behind evolution and excellence and will reveal analogies and surprising differences between two worlds that are so distant, yet so intimately linked. The exhibition aims to emphasise the importance of multidisciplinary approaches in scientific research and invites the exploration of topics that contribute to a deeper understanding of our world.

COMMUNITY TERRITORIES: 
THE VALUE OF COOPERATION

The exhibition explores the deep connection between landscape and community, highlighting how the territory was not only shaped for practical reasons but also for symbolic and social purposes. 

In fact, Friuli's mounds and hillforts are not only archaeological evidence but also convey an idea of community in which cooperation plays a crucial role. An example of this is the construction of the Udine Castle hill, which provides evidence of the importance of planning and sharing resources and skills. 

This timeless message of cooperation is what inspires the proposal of the Udine Castle hill and the mounds and hillforts of Friuli as candidates for UNESCO World Heritage Site status.

Sant’Osvaldo, funerary mound before excavation (Laboratory of Protohistory, University of Udine)
Sant’Osvaldo, 2000 excavation campaign (Laboratory of Protohistory, University of Udine)

MEMORY IN THE LANDSCAPE

During the Early Bronze Age, the upper Friulian plain was dotted with dozens of burial mounds, distributed over a ~10-km-wide strip of land, between the Torre and Tagliamento rivers, and a few fortified settlements (hillforts or ‘castellieri’).

The burial mounds are artificial earthworks 25-30 metres in diameter and about 6.5-7 metres high, which are now preserved in limited numbers. They housed the remains of prominent members of Early Bronze Age communities and date back to the beginning of the 2nd millennium BC, i.e. to a time before the hillforts were built. In some cases, these burial mounds maintained their relevance for centuries, taking on different meanings and functions: between the 19th and 13th centuries BC, in addition to their funerary role, they represented symbols of territorial control by the communities that had erected them; a function that would later be taken over by the fortified settlements. They were also places of congregation and celebration, lookouts and landmarks in exchange routes.

The Friulian hillforts developed at a slightly later stage, spreading mainly during the Middle Bronze Age. An exception is the hillfort of Sedegliano, where five tombs dating between 1900 and 1600 BC were found within the settlement ramparts. The blooming of the hillforts dates back to the Middle Bronze Age, with the building of the castellieri of Udine, Variano di Basiliano, Galleriano di Lestizza and Savalons di Mereto di Tomba. At later stages, other hillforts were built along the springs belt and near rivers. Between the end of the Middle Bronze Age and the beginning of the Recent Bronze Age (14th-13th centuries BC) there was a widespread occupation of the lower Friulian plain.

CONNECTIONS

The spread of the hillforts seems to have been influenced by the presence of the old burial mounds, often placed in intermediate positions between one hillfort and the next. This leads to think that the mounds may have marked the boundaries between the territories of the different communities. The distribution of the early settlements (Middle Bronze Age) seems to suggest a clear awareness of - and perhaps even respect for - the territory of each community, as the settlements were built at regular distances ranging between 8 and 10 kilometres.

General distribution of hillforts and mounds in Friuli. The Udine site is highlighted by a red square (from Pizziolo 2024)

MILLENNIAL UDINE

Based on archaeological findings, the protohistoric settlement of Udine extended south and west of the Castle hill, from the present-day via Mercatovecchio to the former civil hospital and from via Manin to the Chiesa di San Francesco.

With a settlement continuity of about one thousand years, from the Middle Bronze Age to the beginning of the Second Iron Age (around 1500-500 BC), it represents the largest and most complex fortified settlement in Friuli's protohistory. Rising on a modest upland, it occupied an area of approximately 20 hectares - five to ten times larger than that of other coeval settlements on the plain - and was surrounded by a rampart and a ditch. The remains are preserved at a shallow depth. The elevated and well-drained position of the hill protected the area from flooding but the protohistoric levels were subjected to erosion and were later altered by medieval urbanisation. 

The structures that emerged include refuse pits, post holes, and pits of different size and function, together with materials such as pottery, spindle whorls, loom weights, the remains of wattle-and-daub walls and burnt clay floors, which yield clues about ancient domestic life. Two dwellings have been identified so far: a house from the Recent Bronze Age, under Palazzo Mantica, with a rectangular floor plan, wooden walls and a hearth that has been renovated several times; another in Piazza Venerio, with an underground storage room for food supplies, destroyed by fire in the 6th- early 5th century BC. A craft area for recasting small bronze objects, active from the Middle Bronze Age to the early Iron Age, was found beneath Palazzo Dorta. No necropolis has been identified with certainty, but some objects - two fibulae from Piazza I Maggio and Planis and an axe found in Viale Ungheria - seem to refer to Iron Age burial contexts that are now lost.

Archaeological contexts uncovered during urban excavations in Udine (Laboratory of Protohistory, University of Udine)

Reconstruction of basal rampart remains (drawing by F. Zendron)
Second rampart build-up: hypothetical construction technique (drawing by F. Zendron)

DEFENDING THE HILL

A short section of the original rampart, preserved to a height of 80 to 120 centimetres, was found underneath Palazzo Mantica on the south-eastern slope of the Castle hill. Although the available data is limited, the defensive wall of Udine must have been a monumental task at the transition between the Middle and Recent Bronze Ages (14th-13th centuries BC). The archaeological excavation revealed three distinct construction phases of the rampart, all characterised by considerable technical expertise. The original core of the wall was first extended using the technique of 'reinforced earth' (it. ‘terra armata’) where wooden structures filled with gravel, clayey silt, and pebbles were arranged in a chequerboard pattern on several levels, sometimes with smaller elements rotated by 45°. The internal wooden structures acted as stabilising barriers and provided solidity and cohesion to the entire construction.

INGENUITY AND COOPERATION

In the advanced stages of rampart construction, lighter structures called 'gabbioni' were used: these were wooden frames filled with materials placed in sacks or baskets. According to an estimate, if 100 workers were engaged each day in moving around 1 cubic metre of soil each, the entire construction would have taken between two and a half and four years.

POWER AND LANDSCAPE

The Castle hill, visible from every corner of the city, rises in the centre of Udine: this is an imposing and isolated presence, some 30 metres high and with a base 200 metres in diameter.

This hill dominates the urban landscape and has for centuries been one of the most recognisable symbols not only of the city, but of the entire region. Legends, folk tales, and fascinating theories have sprung up around it, intriguing scholars of all ages. The most recent archaeological and geo-archaeological investigations (2020-2022) shed light on its nature: it is an entirely man-made structure with overlapping layers of gravel and clay, stabilised by the use of wooden structures. A monumental mound estimated to contain between 400,000 and 450,000 cubic metres of sediment.

Stratigraphic profile of Udine Castle Hill (modified from Fontana et al. 2023)

Its construction, according to available dating, dates back to between the end of the Middle Bronze Age and the beginning of the Recent Bronze Age (around 1400-1300 BC). The construction of such a majestic artificial hill seems to result from a unified effort: the excavation of the area of the current Piazza I Maggio would have provided the material to build the hill, at the same time generating a reservoir to serve the proto-historic settlement of Udine. A planned work that resulted in a strategic lookout and control of the territory, erected as tangible evidence of the community's power.

COMMUNITY PROJECT

The review of the most recent geo-archaeological data concerning the Udine Castle hill highlighted the most relevant aspects of an extraordinarily planned project, the implementation of which required the organised cooperation of an entire community.
Between 400,000 and 450,000 cubic metres of sediment were moved to shape this imposing upland - a structure in the shape of a truncated cone with terraced sides - mainly taken from the area where Piazza I Maggio is today. The monumental size and complexity of the work provide evidence of a careful planning of the construction phases, the use of established techniques, and the presence of specialised labour engaged over a long period. According to current estimates, with the dedicated labour of 100 workers, the construction would have taken about 10-12 years; with 300 workers 3-4 years; while with 500 workers it could have been completed in just over 2 years (between 26 and 30 months).

To shape the hill

450.000 m³

of debris was moved

Estimated time required / estimated workforce

10 years

100 workers

Estimated time required / estimated workforce

2 years

500 workers

Bronze Age communities
comprised around

100 individuals

Daily earth handling
using baskets

1 m³

1700 AD

1500 m³ 

of debris moved

in 32 days

by 48 workers

COOPERATION

Given the small size of Friuli's Bronze Age communities - each consisting of a few hundred people - it is plausible that the construction of Udine's Castle hill required the contribution of labour from neighbouring settlements. In a pre-industrial society, equipped only with tools made of wood and horn, and simple baskets, it is estimated that each worker could have moved around 1 cubic metre of soil per day. This theoretical figure is supported by more recent data: in 1700, during the reclamation of the pond that occupied what is now Piazza I Maggio (then known as Giardin Grande), 48 workers equipped with metal shovels took 32 days to remove 1,500 cubic metres of soil, an average daily rate per person surprisingly similar to that estimated for the protohistoric age.

THE ALTRUISM DILEMMA.
WHAT IS THE FUNCTION OF ALTRUISM IN NATURE?

For human beings, altruism is a concept that brings with it ethical implications and philosophical questions: an action is altruistic if it is carried out with the conscious intention of helping someone else.

Does something similar exist in other species? Altruism in the animal world is still being debated by scientists and remains partly shrouded in mystery. The phenomenon is difficult to define and apparently contradicts the very pillars of the theory of evolution. It was while studying the social systems of the Hymenoptera (bees, wasps and ants) that Charles Darwin was confronted with a dilemma that threatened to undermine his theory. The presence among insects of entire “altruistic” castes that renounce their own offspring to devote their lives to the common good was at odds with the principle of natural selection of the fittest, if by “fittest” we mean the individual with the greatest reproductive success. According to Darwin, this dilemma posed crucial questions for science, going beyond insects to involve even the most complex of societies, that of human beings. 

'He who was ready to sacrifice his life, rather than betray his comrades, would often leave no offspring to inherit his noble nature'.

from The Descent of Man and Selection in Relation to Sex, 1871

Altruism can be an effective choice with many benefits and it is by no means a prerogative of the human species. On the contrary, mammals, birds, fish, even invertebrates and microorganisms also “know” how to be altruistic.

STRUGGLE OR COOPERATION?

What is the difference between a lone wolf and a wolf living in a pack?

We tend to think of species with complex social behaviour as somehow being more highly evolved.  It is not by chance that we consider ourselves to be the triumph of evolution! In reality, social life involves many costs. Animals living in groups must compete with their peers for space, food and partners and are more exposed to the risk of epidemics.

So are competition and the struggle for survival the only drivers of evolution?

The Russian philosopher and zoologist Pyotr A. Kropotkin believed that mutual support was as much a law of animal life as fighting each other and that it was even more important because of the benefits it brought to the species practising it. He wrote:

'Wherever I saw animal life in abundance, as, for instance, on the lakes where scores of species and millions of individuals came together to rear their progeny; in the colonies of rodents; in the migrations of birds which took place at that time along the Usuri […]: in all these scenes of animal life which passed before my eyes I saw Mutual Aid and Mutual Support'.

Mutual Aid - A Factor of Evolution, 1902

By living together, animals can, in fact, help each other in many ways.

The choice to live a social life is thus based on a continuous balance between costs and benefits. It’s all about equilibrium. When the individuals derive an immediate benefit from a behaviour, we speak of cooperation such as in hunting or defence against predators. On the other hand, when individuals behave altruistically because they know they will receive a reward later on, this is referred to as reciprocity, for example, in social grooming when two individuals reciprocate to eliminate each other’s parasites. In other words, the maxim “do as you would be done by” or, more cynically, “nobody does anything for nothing” also applies in the animal world.

FIRST AID

Matabele ants (Megaponera analis) rescue their wounded comrades on the “battlefield”. They transport them to the safety of the anthill where they treat them by lengthy licking. The treatment is reserved for ants with a good chance of survival.

THE MUTUAL AID

Duels, battles and even wars, such as those between ants and termites: in nature, there is no shortage of examples where competition between individuals and between species becomes ruthless

But certain types of interaction characterised by cooperation, or at least mutual tolerance, also exist between different species, in which the species involved "barter" with each other resources or "services. These relationships may be occasional, long-lasting or even indispensable for the survival of the organisms involved, as in obligate symbioses, when two species are strictly interdependent. The life of one would be impossible without the life of the other!

When the species involved in the interaction benefit equally from the relationship, we speak of “mutualism”. A classic example is the alliance between plants and the insects that guarantee pollination as they feed on the nectar produced by the flowers. But inter-species solidarity can take the most varied forms, with each species contributing its own strong points to the life of the community. The alarm calls of birds that reptiles have learned to recognise. 

The joint ventures of predators, each with their own hunting technique. Skin parasites as annoying to pachyderms as they are delicious for cleaner birds. The huge bodies of sharks that act as taxis for harmless pilot fish.

The network of more or less opportunistic, more or less advantageous, exchanges between living beings is part of the great interconnected system known as ecology.

JOINT VENTURE

A cooperative hunting coalition between the celestial grouper (Cephalopholis argus) and the whitemouth moray (Gymnothorax meleagris). To engage the moray in joint hunting, groupers employ visual signals: they call the morays from their burrows by shaking their heads until the morays follow. When hunting together on the reef, the two species combine their complementary techniques: the grouper positions itself near an opening to block the prey’s escape route, while the moray eel moves inward. In this way, the prey is effectively trapped, and both species benefit from increased hunting success.

COSTS AND BENEFITS

When we look at altruism in the animal world through a human lens, we risk anthropomorphising animal behaviour, attributing it with moral significance.

A biologist, however, weighs behaviour differently: an action is considered truly altruistic only if its costs outweigh its benefits. In some species, individuals appear to make radical choices, such as giving up reproduction or even sacrificing their own lives. Why do they do this? It is the same question Charles Darwin asked when reflecting on the distinct social roles within insect colonies, roles that shape the lives and destinies of individual members:

‘This difficulty, though appearing insuperable, is lessened, or, as I believe, disappears, when it is remembered that selection may be applied to the family, as well as to the individual, and may thus gain the desired end.’

C. Darwin, On the Origin of Species, 1859

The key to understanding altruistic behaviours therefore appears to lie in the genetic heritage shared among relatives.

But is it possible to extend the concept of family to groups of individuals who are not closely related by kinship but are nonetheless cohesive?

If we imagine a situation in which, within a species, many groups compete with each other for resources, it is advantageous within the group to be altruistic, to cooperate and to help one another. This theory, known as group selection, is able to explain the most radical forms of altruism, those expressed beyond the family.

HELPERS IN THE NEST

In many species of birds and mammals, some individuals forgo reproduction or delay it by a few years to help their parents raise their siblings, providing food and protecting the young from predators. The decision whether or not to become a helper can depend on the availability of quality habitat or reproductive partners. If these are scarce, it may be advantageous to ensure the survival of one’s siblings, with whom an individual shares 50% of their genetic makeup.

THE MISTERY OF EMOTIONS

Cooperation and reciprocity do not, however, explain all the altruism phenomena occurring in nature. Something seems to break the logic of opportunism or ties between blood relatives.

Scientists have sought new evolutionary explanations that interpret natural selection in a more complex way, analysing multiple factors. One of these that helps explain the significance of altruistic behaviour is empathy, an emotional involvement that drives social animals to go beyond the cold cost-benefit calculation of their actions and that can also manifest itself between individuals of different species.

In some animals, empathy - the ability of an individual to recognise the moods of others and put itself “in their shoes” - gives rise to acts of selfless altruism that are no longer the prerogative of humans alone. Helping those in distress, consoling those going through a difficult time, even mourning their dead as do elephants, are all documented behaviours in the animal kingdom. 

These reactions to the plight of fellow beings might seem mysterious or inexplicable, were it not for the fact that they mirror exactly what happens amongst us humans.

FRIENDSHIP OR CHOCOLATE?

Rats suffer stress when they see their companion imprisoned. Laboratory experiments on the empathic nature of their behaviour have shown that these rodents prefer to free their fellow rats and share a bar of chocolate with them, rather than selfishly keep all the food for themselves, with a “lump in their throat”.

WE ALL HATED EACH OTHER SO MUCH

After a conflict between two chimpanzees (Pan troglodytes), the other members of the group often decide to console the one who has had the worst of it with kisses and caresses. It is still not entirely clear why they do not decide to flatter the winner instead, a choice they might find more advantageous.  At the end of the fight, the protagonists also often seek reconciliation by exchanging similar gestures of affection. When they fail to do so spontaneously, a female mediates. These behaviours restore harmony and contribute to the well-being of the whole group.

THE ENEMY OF MY ENEMY IS MY FRIEND

Humpback whales (Megaptera novaeangliae) defend animals of other species from attacks by killer whales (Orcinus orca) by interfering with their hunting manoeuvres. It is unclear why they risk injury and waste so much energy protecting other cetaceans, pinnipeds and even sunfish. Or perhaps their aim is just to harm one of their most bitter enemies? Killer whale attacks are unlikely to bring down the huge adult humpback whale, but may, in fact, be fatal to their offspring.

ONGOING DISCOVERY

An exhibition is an opportunity to explore new themes and stimulate curiosity. But it does not stop there.

It is also a unique opportunity to delve deeper. It can become an essential tool for collecting verifiable and up-to-date data and, finally, it can help develop new knowledge, involving - thanks to books and popular articles and videos - readers of all ages and levels of knowledge. In this process, the Museum naturally plays a key role and, in collaboration with people engaged in research and its dissemination, can offer valuable insights. Reading and reflecting are indeed simple acts, but they can produce major changes, since knowledge grows when it is shared.

The scientific articles that inspired this exhibition

EMPATHY

• Ben-Ami Bartal I., Decety J., Mason P. 2011 - Empathy and Pro-Social Behavior in Rats, Science, 34(6061): 1427-1430. Link
• De Waal F.B. M. 2008 - Putting the Altruism Back into Altruism: The Evolution of Empathy, Annual Review of Psychology, 59: 279-300. Link
• De Waal F.B. M., Aureli F. 1999 - La risoluzione dei conflitti nei primati, in Enciclopedia Treccani, Frontiere della Vita. Link
• Frank E.T., Wehrhahn M., Linsenmair K. E. 2018 - Wound treatment and selective help in a termite-hunting ant, Proceedings of the Royal Society. Biological sciences, 285(1872): 20172457. Link
• Miralles A., Raymond M., Lecointre G. 2019 - Empathy and compassion toward other species decrease with evolutionary divergence time, Scientific Reports, 9: 19555. Link
• Pérez-Manrique A., Gomila A. 2018 - The comparative study of empathy: sympathetic concern and empathic perspective-taking in non-human animals, Biological Reviews. Cambridge Philosophical Society, 93(1): 248-269. Link
• Zahn-Waxler C., Hollenbeck B., Radke-Yarro M. 1984 - The Origins of Empathy and Altruism, in Fox M. W., Mickley L. D. (eds), Advances in animal welfare science 1984/85, Washington DC, The Humane Society of the United States: 21-41. Link

ALTRUISM

• Dunbar R. 1991 - Functional Significance of Social Grooming in Primates, Folia Primatologica, 57(3): 121-131. Link
• Nazzi F., Vianello A. (eds) 2018 - L'altruismo. Competizione e cooperazione dalla biologia all'economia, dalla filosofia alle neuroscienze, Quaderni di Aperture. Idee, scienza e cultura, Udine, Forum Edizioni. Link
• Picard A., Mundry R., Auersperg A., Boeving E., Boucherie P., Bugnyar T., Dufour V., Emery N., Federspiel I., Gajdon G., Guéry J.-P., Hegedič M., Horn L., Kavanagh E., Lambert M., Massen J., Rodrigues M., Schiestl M., Schwing R., Szabo B., Taylor A., van Horik J., von Bayern A., Seed A., Slocombe K. 2019 - Why preen others? Predictors of allopreening in parrots and corvids and comparisons to grooming in great apes, Ethology, 126: 10.1111/eth.12999. Link
• Pievani T. 2010 - Biologia dell’altruismo, Micromega, Almanacco della scienza, 7: 45-63. Link
• Pitman R. L., Volker B. D., Gabriele C. M., Srinivasan M., Black N., Denkinger J., Durban J. W., Mathews E. A., Matkin D. R., Neilson J. L., Schulman-Janiger A., Shearwater D., Stap P., Ternullo R. 2017 - Humpback whales interfering when mammal-eating killer whales attack other species: Mobbing behavior and interspecific altruism?, Marine Mammal Science, 33: 7-58. Link
• Strassmann J. E.,  Zhu Y., Queller D. C. 2000 - Altruism and social cheating in the social amoeba Dictyostelium discoideum, Nature 408(6815): 965-967. Link
• Carter G. G., Wilkinson G. S. 2015 - Social benefits of non-kin food sharing by female vampire bats, Proceedings of the Royal Society, Biological sciences, 282: 20152524. Link
• Carzon P., Delfour F., Dudzinski K., Oremus M., Clua É. 2019 - Cross-genus adoption in delphinids: One example with taxonomic discussion, Ethnology, 125(9): 669-676. Link
• González-Forero M., Peña J. 2021 - Eusociality through conflict dissolution, Proceedings of the Royal Society. Biological sciences, 288: 20210386. Link
• Mittal D., Chakrabarti S., Khambda S. B., Bump J. K. 2020 - Spots and manes: the curious case of foster care between two competing felids, Ecosphere Naturalist, 11(2): e03047. Link
• Razik I., Brown B. K. G., Page R. A., Carter G. G. 2021 - Non-kin adoption in the common vampire bat, Royal Society Open Science, 8: 201927. Link
• Rosenbaum S., Gettler L. T. 2018 - With a little help from her friends (and family) part I: the ecology and evolution of non-maternal care in mammals, Physiology & Behavior, 193(part A): 1-11. Link
• Scornavacca D., Cotza A., Lovari S., Ferretti F. 2018 - Suckling behaviour and allonursing in the Apennine chamois, Ethology Ecology & Evolution, 30(5): 1-14. Link
• Wilson A. D. M., Krause J. 2013 - Repeated Non-Agonistic Interactions Between a Bottlenose Dolphin (Tursiops truncatus) and Sperm Whales (Physeter macrocephalus) in Azorean Waters, Aquatic Mammals, 39(1): 89-96. Link

COOPERATION

• Blumstein D. T. 2007 - The Evolution. Function and Meaning of Marmot Alarm Communication, Advances in the Study of Behavior, 37: 371-401. Link
• Clutton-Brock T., O`Riain J. M., Brotherton P., Gaynor D., Kansky R., Griffin A., Manser M. 1999 - Selfish sentinels in cooperative mammals, Science, 284(5420): 1640-1644. Link
• Fontana A., Vinci G., Ronchi L., Mocchiutti A., Muscio G., Visentini P., Bassetti M., Novellino M. D., Badino F., Musina G., Bonomi S.  2023 -The largest prehistoric mound in Europe is the Bronze-Age Hill of Udine (Italy) and legend linked its origin to Attila the Hun, Scientific Report, 13(1): 8848. Link
• Kropotkin P. 2020 - Il mutuo appoggio. Un fattore dell’evoluzione, Milano, elèuthera (trad. it., Mutual Aid. A factor of evolution, 1902, New York, McClure Phillips & Co.). Link
• Lenti Boero D. E. 1992 - Alarm calling in Alpine marmot (Marmota marmota L.): evidence for semantic communication, Ethology Ecology & Evolution, 4(2): 125-138. Link
• Vail A. L., Manica A., Bshary R. 2013 - Referential gestures in fish collaborative hunting, Nature Communications, 4(1): 1765. Link
• Vinci G., Vanzani F. 2025 - Bronze Age monumental earthworks of the Friuli plain (NE Italy): from LiDAR-based morphometric analysis to the reconstruction of settlement patterns and organisation, Archaeological and Anthropological Sciences, 17: 19. Link
• Visentini P. (ed.) 2023 - Archeologia Urbana a Udine. Contributi per una rilettura dei dati provenienti dal colle del Castello, Udine, Comune di Udine, Edizioni del Museo Friulano di Storia Naturale, 58, 398 p. Link
• Visentini P. (ed.) 2024 - Archeologia Urbana a Udine, Udine, Comune di Udine, Pubblicazioni varie del Museo Friulano di Storia Naturale, 76, 416 p. Link

Recommended readings curated by CLUF Bookshop, Udine, exploring themes related to, but beyond, those addressed in the exhibition ‘Cooperare e distinguersi

COOPERATION

• Emanuela Borgnino, Ecologie native, elèuthera, 2022
• Michael Tomasello, Altruisti nati, Bollati Boringhieri, n. ed. 2025
• Vittorio Lingiardi, Io, tu, noi, Utet, 2019
• Aldo Bonomi (a cura di), Sul confine del margine. Tracce di comunità in itinere, Derive Approdi, 2024
• Adele Clarke, Donna Haraway, Making Kin. Fare parentele, non popolazioni, Derive Approdi, 2022
• Adriana Giannini, Lynn Margulis, La scoperta dell’evoluzione come cooperazione, L’Asino d’oro, 2021
• Stefano Bocchi, L’ospite imperfetto, Carocci, 2021
• Tim Ingold, Siamo linee, Treccani, n. ed. 2024
• Frédéric Maupomé, Muri, Nomos, 2025
• Mac Barnett illustrato da Jon Klassen, Sam e Dave scavano una buca, Terre di Mezzo, rist. 2023
• Marcos Farina, Io, tu e gli altri, Ukids, 2025
• Satomi Ichikawa, Amici, Orecchio Acerbo, 2023
• Anne Jankéliowitch, Isabelle Simler, Regni minuscoli, 2022

CREATION

• Francesco Faccin, Zattere, Corraini, 2024
• Elena Granata, Placemaker, Einaudi, 2021
• Pedro Torrijos, Territori improbabili, Saggiatore, 2024
• Dan Nott, Sistemi nascosti, Quinto Quarto, 2024
• Andrea Staid, Dare forme al mondo, Utet, 2025
• Tim Ingold, Making, R. Cortina, 2019
• Rompere le regole, Utet, 2019
• Bruno Munari, Pensare confonde le idee, Corraini, 2023
• Michele De Lucchi, L’architetto, Corraini, 2024
• Cruschiform, L’odissea dei semi, L’Ippocampo, 2024
• Isabelle Simler, Casa, L’Ippocampo, 2023
• Francesco Spampinato, Irene Rinaldi, Facciamo presente, Topipittori, 2025
• Emmanuelle Figueras, Claire De Gastold, Animali tuttofare, L’Ippocampo, 2022
• Guia Risari, Alessandro Sanna, La Terra respira, Lapis, 2022
• Julie Bernard, Il banchiere di semi, Terre di Mezzo, 2024
• Aude Le Pichon, Arnaud Nebbache, Da capogiro, Donzelli, 2024
• Judith Schlansky, Inventario di alcune cose perdute, Nottetempo, 2020

EVOLUTION

• Rebecca Solnit, Storia del camminare, Ponte alle Grazie, 2022
• David Quammen, L’evoluzionista riluttante, R. Cortina, 2025
• Sean B. Carroll, Una serie di fortunati eventi, Codice ed., 2022
• Gabriele Ferrari, Polvere e ossa, Codice ed., 2023
• Stephen Jay Gould, Questa idea della vita, Codice ed., 2022
• Marco Ferrari, L’evoluzione è ovunque, Codice ed., 2021
• Christiane Vadnais, Faune, Codice ed., 2023
• Henry Bergson, L’evoluzione creatrice, Bur Rizzoli, rist. 2025
• Mauro Garofalo, La tigre e l’usignolo, Nottetempo, 2025
• Stephen Jay Gould, Il pollice del panda, Saggiatore, 2016
• Giorgio Manzi, Antenati, Il Mulino, 2024
• Fabien Grolleau, Thomas Brochard-Castex, Grande oceano, Add ed., 2023
• Nicoby Vincent Zabus, Il mondo di Sofia, Longanesi, 2022
• Roberta Ragona Tostoini, Fossili viventi, Aboca kids, 2025
• Serenella Quarello, Alessio Alcini, Estintopedia, Camelozampa, rist. 2024
• Aina Bestard, Paesaggi perduti della Terra, L’Ippocampo, 2020
• Anna Claybourne, Wesley Rbins, Meravigliosa evoluzione, Editoriale Scienza, 2020
• Desmond Morris, La scimmietta nuda, Bompiani, 2025
• Neil Packer, Unico nel suo genere, Camelozampa, 2022
• David Graeber, David Wengrow, L’alba di tutto, Bur Rizzoli, rist. 2025
• Ingannare il tempo. Bruno Munari archeologo, Corraini, 2024

INTERCONNECTIONS

• Venezia e l’Antropocene, Wetlands, 2022
• Serge Latouche, Il disastro urbano e la crisi dell’arte contemporanea, elèuthera, 2025
• Fred Bodsworth, L’ultimo dei chiurli, Adelphi, 2025
• Anna Marson, Antonella Tarpino, Sguardi sul paesaggio, Manifestolibri, 2025
• Niccolò Scaffai, Sotto l’inesauribile superficie delle cose, Aboca, 2025
• Marco Deriu, Rigenerazione, Castelvecchi, 2022
• Donna Haraway, Chthulucene, Nero ed., 2023
• Tim Ingold, Il futuro alle spalle, Meltemi, 2024
• Andrea Staid, Essere natura, Utet, 2022
• Valentina Gottardi, Lumen, Cocaibooks, 2024
• Adriano Favole, Vie di fuga, Utet, 2021

HUMANITY AND NATURE

• Long Litt Woon, La via del bosco, Iperborea, 2023
• Alberto Moravia, Storie della preistoria, Bompiani, 2017
• Giorgio Vallortigara, Pensieri della mosca con la testa storta, Adelphi, 2021
• Giorgio Vallortigara, Altre menti, Il Mulino, 2000
• Giorgio Vallortigara, Cervello di gallina, Bollati Boringhieri, 2024
• Sy Montgomery, Il tempo delle tartarughe, Aboca, 2025
• Paco Calvo, Planta Sapiens, Saggiatore, 2022
• Francesca Buoninconti, Senti chi parla, Codice ed., 2021
• Nicholas P. Money, Natura veloce, natura lenta, Codice ed., 2022
• Eva Meijer, Linguaggi animali, Nottetempo, 2021
• Emmanuelle Pouydebat, Quando gli animali e le piante ci ispirano, Espress, 2021
• Emmanuelle Pouydebat, L’intelligenza animale, Corbaccio, 2018
• Telmo Pievani, La natura è più grande di noi, Solferino, 2022
• Umani e non umani. Noi siamo natura, Utet, 2024
• Aimee Nezhukumatathil, Un mondo di meraviglie, Nottetempo, 2024
• Francesco Remotti, Fare umanità, Laterza, 2013
• Nat Cardozo, Origine, L’Ippocampo 2023

BEYOND BARRIERS

• Bell Hooks, Scrivere oltre la razza, Saggiatore, 2024
• Alexis Pauline Gumbs, Undrowed, Timeo, 2023
• Lorraine Daston, Contro natura, Timeo, 2024
• Lino Leonardi, Razza, Il Mulino, 2024
• Mirella Orsi, Sergio Ferraris (a cura di), Prime. Dieci scienziate per l’ambiente, Codice ed., 2023
• Guido Barbujani, L’alba della storia, Laterza, 2024
• Jackie Higgins, Senzienti, Longanesi, 2025
• Tim Ingold, Antropologia, Meltemi, 2020
• Nicole Shukin, Capitale animale, Tamu, 2023
• Annalisa Strada, Mary Anning, La cacciatrice di fossili, Editoriale scienza, 2025
• Cristina Petit, Qualcosa che c’entra con la felicità, Pulce, 2021
• Issa Watanabe, Migranti, Logos, 2023
• Federica Buglioni, Alfabeti naturali, Topipittori, 2023
• La biblioteca degli animali di Tatsu Nagata, L’elefante, Nomos, 2025
• La biblioteca degli animali di Tatsu Nagata, L’ape, Nomos, 2025
• Rosie Haing, Noi animali umani, Donzelli, 2022
• Chiara Grasso, Una famiglia bestiale, Macadamia, 2024